Family Scarabaeidae

The scarab beetles

reviewed by H.F. Howden, October 1995

Subfamily Scarabaeinae

Tribe Scarabaeini

Boreocanthon Halffter

• B. depressipennis (LeConte), TX-ND-NC-FL (ALA, BAK, CAL, CLA, DIX, DUV, GAD, HER, HIG, HIL, LEV, LIB, MARI, PIN, PUT, TAY, VOL, WAL)
• B. probus (Germar), AZ-UT-NJ-FL (ALA, BRO, DAD, DIX, HIG, HIL, IND, JEF, LAK, LEE, LEO, LEV, ORA, PIN, POL, PUT, SLU, VOL)

Canthon Hoffmansegg

• C. (C.) chalcites (Haldeman), TX-NB-NM-FL (ALA, DAD, FLA, LIB, MARI, PUT, VOL)
• C. (C.) pilularius (Linneaus), TX-MT-MA-FL (ALA, BAK, BRE, CIT, COLU, DAD, DIX, GLA, HAR, HEN, HIL, IND, JAC, LAK, LEV, LIB, MAN, MARI, NAS, PIN, PUT, SAR, SEM, SUW, TAY, VOL)
• C. (C.) vigilans LeConte, TX-KS-MA-FL (ALA, BRA, ESC, GAD, HER, JEF, LEV, LIB, MARI, OKA, PAS, SRO)
• C. (Glaphyrocanthon) viridis viridis (Beauvois), TX-MI-NY-FL (ALA, DAD, DIX, IND, LAK, LEO, LIB, MARI, MON, VOL), Mexico

Deltochilum Eschscholtz

• D. gibbosum gibbosum (Fabricius), TX-KY-NC-FL (ALA, DAD, FRA, HIG, IND, JEF, LEO, LEV, LIB, MARI, MON, OKA, ORA, PIN, PUT, SEM, VOL)

Melanocanthon Halffter

• M. bispinatus (Robinson), MS-NJ-FL (ALA, BAK, CAL, COLU, DIX, FRA, GAD, LEO, LEV, LIB, VOL)
• M. granulifer (Schmidt), TX-FL (ALA, BAK, CIT, CLA, COLU, DIX, FRA, HAM, JEF, LAK, LEV, MAN, MARI, PAS, PUT, SUM, SUW)
• M. punctaticollis (Schaeffer), FL endemic (ALA, COLU, DES, HIG, HIL, LEV, MARI, PIN, PUT)

Pseudocanthon Bates

• P. perplexus (LeConte), AZ-IL-NC-FL (ALA, BAY, CHA, COLL, COLU, DAD, DUV, HIG, IND, JAC, LAK, LEE, LEO, LEV, LIB, MAN, MON, OKA, PIN, SAR), Mexico, CAmer, SAmer

Tribe Coprini

Ateuchus Weber

• A. histeroides histeroides Weber, TX-NJ-FL (ALA, JAC, LIB); rare
• A. lecontei (Harold), AL-NJ-FL (ALA, BAK, BAY, BRO, CAL, CHA, CLA, DIX, DUV, FRA, GAD, GIL, HER, HIG, HIL, IND, JAC, JEF, LAK, LEO, LEV, LIB, MARI, MON, NAS, OKA, ORA, OSC, PAS, PIN, POL, PUT, SJO, SRO, SAR, SEM, TAY, VOL)

Copris Müller

• C. gopheri Hubbard, FL endemic (ALA, GIL, HIG, LEV, PIN, SEM); in gopher tortoise burrows; threatened
• C. howdeni Matthews and Halffter, FL endemic (IND, MAN, MON, POL) (Woodruff & Deyrup 1995 apparently do not accept Lampert's 1977 South Carolina record); rare
• C. inemarginatus Blatchley, FL endemic (ALA, CLA, DAD, DIX, DUV, GIL, HAR, HIG, HIL, LEV, MARI, PIN, PUT, SEM, VOL)
• C. minutus (Drury), TX-IA-NH-FL (ALA, BAK, BAY, CHA, CLA, COLL, DES, DIX, DUV, FLA, GAD, GIL, GLA, HAM, HAR, HEN, HER, HIG, HIL, IND, JAC, JEF, LAF, LEO, LEV, LIB, MARI, OKA, ORA, OSC, PIN, PUT, SAR, SEM, SUM, VOL, WAK)

Dichotomius Hope

• D. c. carolinus (Linnaeus), TX-SD-MA-FL (ALA, CHA, DAD, GAD, HIL, JAC, JEF, LEO, LIB, MARI, OKA, ORA, SRO, SEM, WAS)

Phanaeus MacLeay (see Edmonds 1994)

• P. (Phanaeus) igneus MacLeay, seUS, FL (ALA, BAK, BRO, CAL, COLL, COLU, DAD, DES, FLA, GIL, HAR, HIG, HIL, IND, JEF, LAK, LEV, LIB, MAD, MARI, NAS, ORA, OSC, PAS, PIN, PUT, SAR, SEM, SJO, SUW, VOL, WAL)
  = P. igneus floridanus d'Olsoufieff
• P. (Phanaeus) vindex MacLeay, TX-SD-MA-FL (ALA, BAK, BRE, CAL, CLA, DAD, DUV, ESC, FLA, GAD, GIL, GLA, GUL, HIG, HIL, JAC, JEF, LAK, LEO, LEV, MAD, MAN, MARI, MON, PAL, POL, PUT, SAR, SEM, SUM, TAY, UNI, VOL)
  = P. vindex cyanellus Robinson
• P. (Phanaeus) t. triangularis (Say), seUS, FL (LIB)
• = P. torrens niger d'Olsoufieff

Tribe Onthophagini

Onthophagus Latreille

• O. aciculatulus Blatchley, FL endemic (HIG, PIN, PAS); status undetermined
• O. concinnus Laporte, MS-TN-PA-FL (ALA, DAD, LEO, LIB)
• +O. depressus Harold, immigrant from Africa, GA-FL (HIG, IND)
• +O. gazella Fabricius, the brown dung beetle, introduced from Africa to aid decomposition of cattle dung, TX-FL (ALA, HIG, LEV, LIB) (see Downie 1984)
• O. hecate blatchleyi Brown, SC-GA-FL endemic (ALA, BAK, BRO, COLL, COLU, DAD, GLA, HAR, HEN, HIG, HIL, IND, JAC, LAK, LEE, LEO, LEV, LIB, MAN, MARI, ORA, OSC, PAL, PIN, POL, PUT, SAR, SEM, SLU, VOL)
• O. oklahomensis Brown, TX-KS-VA-FL (ALA, BAK, CAL, CLA, DIX, ESC, HAR, HER, HIG, HIL, LAF, LAK, LEE, LEO, LEV, LIB, MAN, MARI, PUT, VOL)
• O. orpheus orpheus (Panzer), TX-WI-PA-FL (COLU, IND, JAC, MON, PUT, SUW, VOL); in pack rat nests
• O. pennsylvanicus Harold, TX-WY-NY-FL (ALA, BAK, COLL, ESC, HIG, HIL, JAC, LAK, LEV, LIB, OKA, ORA, POL, PUT)
• O. polyphemi polyphemi Harold, SC-FL (ALA, GIL, LEO, LEV, MARI, PUT); in gopher tortoise burrows; threatened
• O. polyphemi sparsisetosus Howden and Cartwright, MS-FL (CAL, LIB, WAL); threatened
• O. striatulus floridanus Blatchley, FL endemic (ALA, BAK, CAL, CLA, DAD, FRA, IND, LAF, LEE, LEO, LEV, LIB, MAN, MARI, PIN, POL, SEM, VOL)
• O. subaeneus (Beauvois), TX-KS-PA-FL (LEO)
• +O. taurus Schreber, introduced from Europe, seUS, FL (ALA, DIX, FLA, TAY)
• O. tuberculifrons Harold, TX-KS-IL-NJ-FL (ALA, BAK, BAY, CAL, CLA, COLL, DAD, DES, DIX, HAR, HIG, HIL, IND, JAC, LAF, LAK, LEO, LEV, LIB, MAN, MARI, MON, OSC, PAL, PAS, PIN, POL, PUT, SLU, SAR, SEM, VOL)

Tribe Oniticellini

Oniticellus Audinet-Serville

• *O. cubiensis Laporte (questionably from Florida) (MON, Key West?), Jamaica, Cuba, Bahamas

Subfamily Aphodiinae

Tribe Aphodiini

Aphodius Illiger

• A. aegrotus Horn, NC-FL(ALA, BRE, DUV, ESC, GUL, HIG, HIL, JAF, LAK, LEV, MARI, OKA, ORA, PIN, PUT, SEM, TAY, WAL); species of special concern
• A. bicolor Say, TX-NB-PQ-FL (ALA, BAK, LEV, PIN)
• A. campestris Blatchley, AL-NJ-FL (ALA, BAK, BAY, BRE, CAL, CLA, COLL, DAD, DUV, ESC, FLA, GAD, GLA, GUL, HAR, HIG, HIL, HOL, IND, JAC, JEF, LAF, LEE, LEO, LEV, LIB, MAN, MARI, NAS, OKA, ORA, OSC, PAS, PIN, POL, PUT, SAR, SEM, SLU, SRO, TAY, VOL, WAS)
• A. crassulus Horn, VA-FL (ALA, BAK, BRA, LEO, LEV, LIB, PAS, PUT)
• A. dyspistus Skelley and Woodruff, AL-FL (ALA, FLA, GIL, LEV, OKA, PUT, WAL); with pocket gophers
• A. fimetarius (Linnaeus), CA-BC-NS-FL (ALA, CLA, DIX, GAD, JAC, JEF, LEV, ORA, PUT)
• A. floridanus Robinson, FL endemic (ALA, BAK, COLL, DAD, FLA, GAD, GLA, HEN, HIG, IND, JAC, JEF, LEO, LEV, MARI, PUT, SAR, SEM, SRO)
• A. fordi Gordon, GA, FL (DIX, new state record)
• +A. granarius (Linnaeus), cosmopolitan, US, FL (JAC, WAL), immigrant from Europe
• +A. haemorrhoidalis (Linnaeus), immigrant from Europe, MO-NB-FL (ALA, BRA, LAK, OSC, PUT)
• A. haldemani Horn, TX-KS, FL (ALA); rare
• A. hubbelli Skelley and Woodruff, AL-FL (ALA, FLA, OKA, PUT, WAL); with pocket gophers; species of special concern
• A. laevigatus Haldeman, AL-GA-FL (ALA, BRE, DUV, FLA, GIL, HIL, LAK, LEV, MARI, ORA, PIN, PUT, SEM, WAL); with pocket gophers; species of special concern
• *A. lentus Horn, IL-MA-GA-?FL (ESC, doubtful record)
• +A. lividus (Olivier), immigrant from Europe, cosmopolitan, US, FL (ALA, BAK, BRO, CAL, CHA, CLA, COLL, DAD, DUV, ESC, FLA, GAD, GLA, GUL, HAR, HIG, HIL, IND, JAC, JEF, LAK, LEE, LEO, LEV, LIB, MAN, MARI, MON, NAS, OKA, OKE, ORA, PAL, PAS, PIN, POL, PUT, SAR, SRO, TAY, VOL, WAS), Cuba to Puerto Rico, Bahamas, Mexico to Argentina
• A. lutulentus Haldeman, TX-IK-OH-FL (ALA, LEO)
• A. nigritus Fabricius, TX-SC-FL (ALA, BAK, CHA, COLL, DAD, GLA, HAR, HIG, HIL, IND, LEE, LIB, MART, MON, OKE, ORA, PAS, PIN, PUT, SAR, SEM, SJO, SUM, TAY, VOL), WInd, Mexico, CAmer, Old World
  = A. cuniculus Chevrolat
• A. parcus Horn, FL endemic (ALA, BAK, BRO, CHA, COLL, DAD, DUV, HIG, IND, LEE, MAN, MARI, PAL, PIN, PUT, SAR, SEM, TAY, VOL)
• A. pholetus Skelley and Woodruff, FL endemic (OKA); with pocket gophers
• A. platypleurus Skelley and Woodruff, AL, FL (LEV, OKA, PUT); with pocket gophers
• A. rubeolus (Beauvois), TX-NB-PQ-FL (ALA, BAK, CAL, ESC, GAD, GUL, HIG, HIL, HOL, JAC, JEF, LEO, LEV, LIB, MAN, MARI, OKA, SRO, TAY, WAS)
• *A. rusicola Melsheimer (questionably from Florida), NM-NB-NY-FL (LEV)
• A. stercorosus Melsheimer, eUS, FL (DAD, ESC, JEF, LEO)
• A. stupidus Horn, AL-NC-FL (ALA, BAK, CAL, GAD, JAC, LEV, LIB, MARI, PUT, WAL)
• A. tanytarsus Skelley and Woodruff, AL, FL (ALA, OKA, WAL); with pocket gophers
• A. troglodytes Hubbard, GA-FL (ALA, DAD, GIL, HIG, IND, LEV, LIB, MARI, PUT, TAY); with gopher tortoise, threatened

Tribe Eupariini

Aphotaenius Cartwright

• A. carolinus (Van Dyke), IN-MD-FL (JAC, LIB, MON); status undetermined

Ataenius Harold (see Cartwright 1974)

• A. abditus (Haldeman), MS-MI-NH-FL (BAK), Mexico?
• A. alternatus (Melsheimer), TX-PA-FL (ALA, BAK, BAY, BRE, BRO, CAL, CHA, CLA, COLL, DAD, DES, DIX, DUV, ESC, GAD, GLA, HIG, HIL, HOL, IND, JEF, LAK, LEE, LEV, MAN, MARI, MON, OKE, ORA, PAL, PAS, PIN, POL, PUT, SAR, SEM, SJO, TAY, VOL, WAS)
• A. apicalis Hinton, TX-NB-PA-FL (LIB)
• A. brevicollis Wollaston, TX-FL (ALA, DAD, MON); rare
• A. cylindrus Horn, TX-NJ-FL (ALA, BAK, BRE, BRO, CHA, CLA, COLL, DAD, DES, DIX, DUV, ESC, FLA, GIL, GLA, HAR, HIG, HIL, IND, JAC, JEF, LEE, LEO, LEV, LIB, MAN, MARI, OKE, PAL, PAS, PIN, POL, PUT, SAR, SEM, SLU, SRO, TAY, WAS)
• A. erratus Fall, TX-OH-FL (ALA, DAD, DUV, GAD, HIG, JEF, LEO, MARI, MON, ORA, SEM, SRO)
• A. exiguus Brown, MS-NC-FL (ALA, BRE, BRO, CLA, COLL, DAD, DUV, FLA, HAR, HER, HIG, HIL, IND, JAC, LEE, LEV, MAN, MARI, MON, PAL, PIN, POL, PUT, VOL)
• A. fattigi Cartwright, TX-MN-MA-FL (ALA, BAK, BRE, BRO, CAL, CHA, CLA, COLL, DIX, DUV, ESC, GLA, GUL, HIG, HIL, HOL, IND, JAC, JEF, LEE, LEO, LEV, LIB, MAN, MARI, OKA, OKE, ORA, PAL, POL, PUT, SAR, SEM, VOL, WAS)
• A. figurator Harold, NM-IN-PA-FL (CAL)
• A. gracilis (Melsheimer), CA-TX-NB-PQ-FL (ALA, BAK, BRO, CAL, CHA, CLA, COLL, DAD, DES, DUV, ESC, GAD, GUL, HEN, HIG, HIL, IND, JAC, JEF, LEE, LEO, LEV, LIB, MAN, MARI, MON, ORA, PAS, PIN, POL, PUT, SAR, SEM, TAY, VOL), Cuba, WInd, Colombia to Argentina
• A. havanensis Balthasar, FL (MON), Barbados, Cuba, St. Croix, St. Kitts; rare
• A. imbricatus (Melsheimer), TX-MN-NH-FL (ALA, BAK, BRE, CAL, CHA, CLA, COLL, DAD, DES, DUV, ESC, FLA, GUL, HIG, HIL, IND, LEE, LEO, MON, OKA, ORA, PAL, PIN, POL, SEM, SRO, TAY, WAS), Mexico, CAmer, Bahamas, Cuba
• A. insculptus Horn, MS-NJ-FL (HIG, ORA, WAL)
• A. integer Harold, FL (ALA, BAY, COLL, DAD, GUL, HIG, HIL, MAN, ORA, POL)
• A. languidus Schmidt, TX-AL-FL (BRO, DAD, HIG)
• A. miamii Cartwright, TX-NJ-FL (ALA, BRO, COLL, DAD, DUV, ESC, FLA, GIL, HIG, IND, LEO, LEV, MON, PAL, POL, WAL)
• A. ovatulus Horn, TX-NJ-FL (ALA, GIL, HIG, LEO, NAS, OSC, WAK)
• A. picinus Harold, TX-SC-FL (ALA, CAL, CLA, DUV, ESC, HOL, JAC, JEF, LEO, LEV, LIB, MON, OKA, PUT, SRO, WAS), Cuba, Jamaica, Hispaniola, Puerto Rico, Lesser Antilles, Venezuela
• A. platensis Blanchard, CA-OK-IN-NC-FL (ALA, BAK, BAY, CAL, COLL, DAD, DUV, ESC, GUL, HAM, HEN, HIG, HIL, HOL, IND, JAC, JEF, LEO, LIB, MAN, MARI, OKA, ORA, POL, SRO, TAY, WAS)
• A. rhyticephalus Chevrolat, TX-SC-FL (ALA, COLL, DAD, DUV, HIG, JEF, LEV, MARI, POL, SAR), Cuba, WInd, Mexico to SAmer
• A. rudellus Fall, FL (BRE, CHA, DAD, DUV, LEE, MON); status undetermined
• A. saramari Cartwright, FL endemic (HIG, LEV, MARI, OSC, SEM, SLU); status undetermined
• A. sciurus Cartwright, FL endemic (ALA, DAD, HIL, LEV, LIB, MAN, PIN, PUT, SUM); in squirrel nests; threatened
  = A. brevinotus Chapin, misidentification
• A. simulator Harold, MS-MD-FL (ALA, BAK, BAY, BRE, CAL, CLA, COLL, DAD, DES, DIX, DUV, ESC, FLA, GAD, GIL, GUL, HAM, HAR, HEN, HIG, HIL, HOL, IND, JAC, JEF, LAK, LEO, LEV, LIB, MAD, MAN, MARI, OKA, OKE, ORA, OSC, PAL, PIN, POL, PUT, SEM, SJO, SLU, SRO, SUW, WAS), SAmer
• A. spretulus (Haldeman), the black turfgrass ataenius, NM-ID-NH-FL (ALA, BRE, BRO, CHA, COLL, DAD, DUV, GLA, GUL, HAR, HIG, HIL, IND, LEE, LEO, MAN, MON, OKE, ORA, PAL, POL, SAR, SEM, SLU)
• A. strigatus (Say), AZ-OR-ON-FL (GUL)
• A. stroheckeri Cartwright, FL (DAD, MAN); status undetermined
• A. superficialis Cartwright, FL (MON); (not a valid species?); status undetermined
• A. wenzelii Horn, TX-IA-PA-FL (FLA, GUL)
• A. woodruffi Cartwright, FL (FLA); status undetermined
• A. sp., FL (DAD)

Euparia Audinet-Serville

• E. castanea Serville, TX-GA-FL (ALA, BAK, BAY, BRA, CHA, CLA, DAD, DES, DUV, GAD, HAR, HER, HIG, HIL, HOL, JEF, LAK, LEE, LEO, LEV, MARI, MON, OKA, ORA, OSC, PAL, PAS, PIN, POL, SEM, TAY, VOL, WAS); in nests of native Solenopsis (see Wojcik 1977, 1978), Mexico, CAmer

Martineziella Chalumeau

• M. excavaticollis (Blanchard), TX-NC-FL (ALA, BAK, BRA, CAL, CHA, CLA, DAD, DUV, ESC, GUL, HER, HIG, HIL, IND, JAC, JEF, LEO, LIB, MARI, OKA, ORA, PIN, POL, SAR, SEM, SRO, SUM, TAY, UNI, WAK, WAS); lives in fire ant nests (Wojcik, 1977, 1978)
  = Myrmecaphodius excavaticollis (Blanchard)

Pseudataenius Brown

• P. contortus Cartwright, LA-NJ-FL (LEO)
• P. socialis (Horn), TX-KS-FL
• +P. waltherhorni (Balthasar), Cuba, LA, PA, FL (DAD, MON), threatened, immigrant?
  = Ataenius walterhorni

Tribe Psammodiini

(see Gordon and Pittino 1992)

Geopsammodius Gordon and Pittino

• G. relictillus (Deyrup and Wooduff), FL endemic (HIG)
• G. hydropicus (Horn), FL (SRO)

Leiopsammodius Rakovic

• L. malkini (Cartwright), FL endemic (ALA, BAK, BRO, CAL, CHA, COLL, DAD, DES, DUV, HIG, IND, LAK, LEE, LEV, MAN, MARI, ORA, OSC, PAL, PIN, POL, PUT, SAR, TAY)

Odontopsammodius Gordon and Pittino
= Psammodius Fallen

• O. armaticeps (Fall), GA-FL (ESC, HIL, JAC, MARI, PIN, SEM)
• 0. bidens (Horn), MS-NJ-FL (CHA, DAD, SJO)
• 0. cruentus (Harold), SC-GA-FL (ALA, DUV)

Platytomus Mulsant
= Pleurophorus Mulsant

• P. atlanticus (Cartwright), (LEV)
• P. caelicollis (Cartwright), FL (ALA)
• P. longulus (Cartwright), TX-FL (ALA, BAK, BRE, BRO, CAL, CHA, COLL, DAD, DUV, ESC, GAD, GUL, HAR, HER, HIG, HIL, HOL, IND, JAC, JEF, LAK, LEE, LEO, LEV, LIB, MAN, MARI, MON, OKA, ORA, PIN, POL, PUT, SEM, SJO, SRO, TAY, WAS), SAmer
• P. micros (Bates), AZ-UT-FL (DAD, BRE), Mexico, Guatemala

Rhyssemus Mulsant

• *R. scaber Haldeman (questionably recorded from Florida), TX-DC-FL (SEM?)

Subfamily Melolonthinae

Tribe Sericini

Serica MacLeay

• S. aemula Dawson, TX, FL
• S. aspera Dawson, AL-NC-FL (ALA, MAR)
• S. atracapilla (Kirby), AL-BC-NS-FL (ALA)
• S. delicata Dawson, FL endemic; status undetermined
• S. floridana Dawson, MS-NC-FL (ALA)
• S. frosti Dawson, FL endemic (HIG); rare
• S. georgiana georgiana Leng, TX-IN-NY-FL
• S. georgiana lecontei Dawson, MS-AL-FL
• S. intermixta Blatchley, NM-MB-PQ-FL (JAC)
• S. iricolor (Say), TN-MA-FL
• S. panda Dawson, AL, IN-FL
• S. parallela Casey, widespread eNAmer, FL (NAS)
• S. peleca Dawson, FL endemic (ALA, LEO)
• S. pusilla Dawson, FL endemic (ALA, LEV, MARI, PIN); status undetermined
  = errans Blatchley
• S. rhypha Dawson, FL endemic (OKA, SRO); status undetermined
• S. sericea (Illiger), widespread NAmer, FL
• S. spicula Dawson, GA-FL
• S. tantula Dawson, FL endemic (IND, PAL); status undetermined
• S. vespertina (Gyllenhall), se US, FL (OKA)

Tribe Liparetrini

Hypotrichia LeConte

• H. spissipes LeConte, FL endemic (ALA, HIG, LAK, LEV, MARI, SJO); uncommonly collected, flies during rain on summer nights; species of special concern.

Tribe Melolonthini

Diplotaxis Kirby (see Vaurie 1960)

• D. bidentata LeConte, NJ-FL (ALA, BRE, CLA, ESC, HIG, IND, LAK, LEV, LIB, MARI, NAS, POL)
• D. frontalis LeConte. GA-FL
• D. languida LeConte, GA-FL (ALA, IND, LEV, MARI, PIN)
• D. liberta (Germar), MS-ON-MA-FL
• D. punctatorugosa Blanchard, TX-RI-FL (ALA, JAC, LIB)
• D. rufa Linell, FL endemic (IND, LAK, MARI); status undetermined
• D. subcostata Blanchard, TX-PA-FL (ALA, LIB, SRO)

Plectris Serville

• +P. aliena Chapin, SC-FL (ALA, NAS); immigrant from SAmer?; a potential turf pest (see Woodruff 1968)

Polylamina Hardy

• P. pubescens (Cartwright), FL endemic (OKA, WAL); rare (formerly in Polyphylla)

Polyphylla Harris

• P. gracilis Horn, FL endemic (LEV, OKA, SRO)
• P. occidentalis (Linnaeus), VA-NC-FL (ALA, LEV, LIB, SRO, SUW)

Phyllophaga Harris (see Woodruff and Beck 1989)

• P. aemula (Horn), TX-NJ-FL (ALA, GAD, JAC, JEF, LEO, LIB, MARI, SRO, WAK)
• P. anxia Leconte, widespread NAmer, FL (LIB); rare
• P. apicata Reinhard, TX-MO-FL (ALA, UNI)
• +P. bruneri Chapin, FL (BRO, COLL, DAD), Cuba, immigrant about 1959
• P. clemens (Horn), TX(?), NJ(?), GA-FL (ALA, LEO); status undetermined
• P. clypeata (Horn), MS-NC-FL (COLL, DIX, DUV, GLA, HEN, HIG, MAN, OKA, OKE, SAR, SEM, SLU, VOL)
• P. crenulata (Froelich), TX-MN-NY-FL (ALA, ESC, GAD, JEF, LEO, LIB)
• P. cupuliformis Langston, TX-SC-FL (ALA, BAK, BAY, BRA, CAL, CLA, COLU, DUV, ESC, FRA, GAD, HIG, HIL, JAC, LAK, LEO, LEV, LIB, MARI, NAS, OKA, ORA, OSC, PAS, PIN, POL, PUT, SRO, UNI)
• P. debilis (LeConte), MS-NC-FL (ALA, BAK, CHA, COLU, DES, DIX, DUV, ESC, FLA, HAM, HIG, HIL, JAC, JEF, LAK, LEO, LIB, MAD, MARI, OKA, ORA, PAS, PUT, SEM, SRO, SUW, TAY, UNI, VOL, WAL, WAS)
• P. diffinis (Blanchard), LA-CO-NJ-FL (ALA, COLU, ESC, GAD, LIB)
• P. dispar (Burmeister), MS-NC-FL (ALA, BAK, BRE, BRO, CAL, CHA, CLA, COLL, COLU, DAD, DUV, ESC, GAD, HAM, HER, HIG, HIL, IND, JEF, LAK, LEE, LEO, LEV, LIB, MARI, OKA, ORA, PIN, POL, PUT, SAR, SEM, SRO, TAY, VOL, WAK)
• P. elizoria Saylor, FL endemic (BRE, HIG, OKE, PAL, POL); rare
• P. elongata (Linell), FL endemic (ALA, BRE, BRO, DAD, GIL, HIG, HIL, LAK, LEV, MARI, ORA, PAL, PAS, PIN, POL, PUT, SAR, SRO, VOL); rare
• P. ephilida (Say), TX-IA-GA-?FL (ALA, DAD, questionable records)
• P. floridana Robinson, FL endemic (ALA, BRE, HER, HIG, HIL, MAN, MARI, PUT, SEM, SJO)
• P. forbesi Glasgow, LA-IL-FL (ESC, SRO); rare
• P. forsteri (Burmeister), TX-WI-NM-FL (DUV, GAD, JAC, JEF, LIB, PUT)
• P. foxii Davis, GA-VA, ?FL (SEM, questionable record)
• P. furtilis (LeConte), TX-MN-MA-FL (LEO)
• P. georgiana Horn, LA-NC-FL (ESC, HAM, JEF, LEO, SRO, SUW)
• P. glaberrima (Blanchard), AL-IL-MA-FL (ALA, BAK, BRE, BRO, CHA, CIT, CLA, COLL, COLU, DAD, DES, DUV, ESC, FLA, GAD, GUL, HAR, HER, HIG, HIL, IND, JAC, JEF, LAK, LEE, LEO, LEV, LIB, MAD, MAN, MARI, MART, MON, OKE, ORA, PAL, PAS, PIN, POL, PUT, SAR, SEM, SJO, SLU, SRO, SUM, SUW, TAY, UNI, VOL, WAK, WAL)
• P. gracilis (Burmeister), LA-IL-MA-FL (ALA, BAK, ESC, HIL, JEF, LEO, LIB, MARI, OKA, SRO, TAY)
• P. hirticula (Knoch), LA-SD-MA-FL (JAC, JEF, LEO, OKA)
• P. hornii (Smith), OK-WI-PA-FL (GAD, LEO, LIB); rare
• P. ilicis (Knoch), TX-ND-MA-GA, ?FL (ALA, questionable record)
• P. implicita (Horn), TX-ND-OH-FL (COLU, GAD, MAD)
• P. infidelis (Horn), AL-GA-FL (ALA, COLU, DIX, HIL, JEF, LEO, LEV, MARI, OKA, PUT, UNI)
• P. knochii (Schönherr and Gyllenhal), MS-KS, IA, MA-FL (ESC, GAD, JAC, JEF, LEO, LIB, SRO)
• P. latifrons (LeConte), MS-NC-FL (ALA, BAK, BAY, BRO, CAL, CHA, COLL, COLU, DAD, DIX, DUV, ESC, FRA, GAD, GLA, GUL, HAM, HAR, HEN, HIG, HIL, HOL, IND, JAC, JEF, LAK, LEE, LEO, MAD, MAN, MARI, MON, OKA, ORA, OSC, PAL, PIN, POL, PUT, SAR, SEM, SLU, SRO, TAY, VOL, WAK)
• P. lota Luginbill, MS-SC-FL (ALA, BAK, CAL, DAD, ESC, JEF, LEO, OKA, TAY, WAK)
• P. luctuosa (Horn), MS-MA-FL (ALA, ESC, GAD, JAC, JEF, LEO, LIB, MARI, OKA, SRO, WAK)
• P. mariana Fall, MS-SC-FL (ALA, ESC, GAD, JAC, LEV, LIB, MARI, OKA, ORA, PUT, SRO, WAL, WAS)
• P. obsoleta (Blanchard), TX-TN-FL (LEO), Mexico, CAmer, SAmer
• P. okeechobea Robinson, FL endemic (HIG, LAK, OKE, POL); rare
• P. ovalis Cartwright, FL endemic (OKA, SRO, WAL); rare
• P. panorpa Sanderson, FL endemic (HIG, POL); rare
• P. parvidens (LeConte), TX-AR-SC-FL (ALA, BAK, BRA, CLA, DES, DIX, DUV, ESC, GIL, HAM, HIL, JAC, JEF, LAK, LEO, LIB, MAD, MAN, MARI, MART, OKA, ORA, PIN, POL, PUT, SEM, SRO, TAY, UNI)
• P. perlonga Davis, MS-IL-FL (GAD, LIB); rare
• P. postrema (Horn), MS-SC-FL (ALA, BAK, BAY, ESC, HIG, HOL, JEF, LIB, OKA, PUT )
• P. profunda (Blanchard), TX-KS-OH-FL (LEO); status undetermined
• P. prununculina (Burmeister), LA-NC-FL (ALA, BAK, BAY, BRE, BRO, CHA, CLA, COLL, COLU, DAD, DUV, ESC, FRA, GAD, GLA, GUL, HAM, HEN, HER, HIG, HIL, IND, JAC, JEF, LAK, LEE, LEO, LEV, LIB, MAD, MAN, MARI, MON, OKA, ORA, OSC, PAL, PAS, PIN, POL, PUT, SEM, SRO, TAY, VOL, WAK, WAS)
• P. pseudofloridana Woodruff and Beck, AL-GA-FL (ALA, BAK, COLU, GAD, HAM, JEF, LEO, LIB, MAD, OKA)
• +P. puberula (Jacquelin du Val), FL (DAD), Cuba, immigrant, not established
• P. quercus (Knoch), LA-KS-PA-FL (ALA, ESC, FRA, GAD, HIL, IND, JAC, JEF, LEO, LEV, LIB, MAD, SRO, WAS)
• P. schaefferi Saylor, LA-GA-FL (COLU, DUV, ESC, GAD, JAC, JEF, LEO, LIB, MAD, NAS, SUW)
• P. skelleyi Woodruff and Beck, FL endemic (ALA, CLA, LAK, LEV, MARI, PUT); rare
• P. subpruinosa (Casey), KS, PA, NC-FL (ALA, HIG, MARI, PAL, PUT)
• P. taxodii Langston, MS-GA-FL (COLU, ESC, LEO, LIB, MAD)
• P. tecta Cartwright, FL endemic (ALA, DUV, HAR, HIG, MAN, MARI, PIN, PUT)
• P. tristis (Fabricius), NM-MN-ME-FL (ESC, LEO, OKA, SRO, WAL)
• P. ulkei (Smith), MS-TN-FL (ALA, ?DUV, ESC, GAD, JAC, JEF, LEO, LIB, MARI, OKA, SRO)
• P. uniformis (Blanchard), AL-NC-FL (ALA, BAK, BRA, CAL, COLU, DAD, DUV, GAD, HOL, JAC, JEF, LEO, LEV, LIB, MAD, MARI, PAS, PUT, SAR, SEM, SUW, TAY, UNI, WAK, WAS)
• P. yemasseei Cartwright, AL-SC-FL (ESC); status undetermined
• P. youngi Cartwright, FL (DAD, only in Alice Wainwright Park at entrance to causeway to Key Biscayne), Bahamas (Andros); rare

Tribe Pachydemini

Gronocarus Schaeffer

• G. autumnalis Schaeffer, AL-FL
• G. multispinosus Howden, FL endemic (Panhandle sand hills); rare; uncommonly collected, males fly during daytime rain in winter; females are flightless.

Tribe Macrodactylini

Macrodactylus Latreille

• M. subspinosus (Fabricius), the rose chafer, IN-VA-GA-FL (LEO)
  = setulosus LeConte

Tribe Hopliini

Hoplia Illiger

• H. equina LeConte, MA-FL
• H. floridana Fisher, FL endemic
• H. meridionalis Boyer, SC-FL

Subfamily Rutelinae

(see Hardy 1991)

Tribe Anomalini

Anomala Samouelle

• A. exigua (Schwarz), FL (Lake Altapopka); rare (extinct?)
• A. eximia Potts, FL (HIG); rare
• A. f. flavipennis Burmeister, eUS, Mexico, FL; rare
• A. f. okaloosensis Potts, FL (DUS, OKA); rare
• A. innuba (Fabricius), eUS, Mexico, FL (DAD, IND, MON)
  = piceola Casey, FL
• A. lucicola Fabricius, eUS, WInd, FL
• A. marginata (Fabricius), eUS, Mexico, FL (DAD, HIG, IND, MARI, MON)
• A. mendica Casey, SC-GA-FL (MAR)
• A. minuta Burmeister, NC-SC-AL-GA-FL
• A. oblivia Horn, the pine chafer, eUS, FL
• A. parvula Burmeister, NC-SC-MS-GA-FL
• A. robinsoni Potts, FL (DAD, MON); status undetermined
  = floridana Robinson
• A. semilivida LeConte, NC-SC-AL-GA-FL
  = flaccida (Casey)
• A. undulata Melsheimer, NC-KY-FL (MON, LAK, PAL)
  = floridana Casey
  = nigropicta Casey
• A. umbra Casey, eUS, FL
  = servilis Casey, FL

Strigoderma Burmeister
= Strigodermella Casey

• S. pygmaea (Fabricius), AL-MO-NJ-FL (ALA, IND, MARI, PAL) (see Bader 1992)

Tribe Rutelini

Cotalpa Burmeister

• C. l. lanigera (Linnaeus), eUS, FL (NAS)

Parastasia Westwood

• P. brevipes LeConte, MO-PA-NY, FL (ALA, LIB, MARI)

Pelidnota MacLeay

• P. punctata (Linnaeus), eUS, FL (ALA, DAD, IND, LEV, MARI, MON)
  = brevicollis Casey, FL (Jacksonville)

Rutela Latreille

• R. formosa Burmeister, FL (DAD, MON), Bahamas, Cuba; status undetermined

Subfamily Dynastinae

Tribe Cyclocephalini

Cyclocephala Latreille (key to South Florida species in Gordon and Anderson 1981.

• C. borealis Arrow, the northern masked chafer, eUS, FL, Mexico
• C. immaculata (Olivier), the southern masked chafer, USA, FL, Mexico, CAmer, SAmer
• C. lurida Bland., FL (ALA, LEO, LEV, LIB, WAL)
• C. miamiensis Howden and Endrodi, FL (DAD, MON); status undetermined; possibly immigrant
• C. parallela Casey, FL endemic (ALA, COLL, DAD, FLA, IND, LAK, MON)
• C. puberula (LeConte), CA-CO-eUS, FL (IND, FLA), Mexico
• C. setidiosa LeConte, FL (OKA)

Dyscinetus Harold

• D. morator (Fabricius), eUS, FL (ALA, LEV, MON), Cuba, Puerto Rico; feeds on water hyacinth (Buckingham and Bennett 1989); often abundant, but seemingly only a minor pest of a few plants (see Woodruff 1970)

Tribe Pentodontini

Aphonus LeConte (sse Gill and Howden 1985)

• A. castaneus (Melsheimer), eUS, FL (ALA, DUV, PIN, SLU, VOL)
• A. variolosus (LeConte), SC-GA-FL (ALA, LEV)

Euetheola Bates

• E. humilis rugiceps (LeConte), the sugarcane beetle, AL-GA-SC-FL; a pest of sugarcane

Ligyrus Burmeister

• L. (Ligyrus) cuniculus (Fabricius), FL, Cuba, Jamaica, Hispaniola, Puerto Rico
• L. (L.) g. gibbosus DeGeer, the carrot beetle, eUS, FL (IND, MON), Cuba, Jamaica, Mexico
  = morio LeConte
• L. (L.) neglectus LeConte, seUS, FL (ALA, HIG, IND, LEV)
• L. (Tomarus) subtropicus (Blatchley), seUS, FL (DAD, MON)
  = blatchleyi Cartwright, FL (Miami)

Tribe Oryctini

Strategus Hope

• S. aloeus julianus Burmeister, sUS, Mexico, CAmer, FL (LIB)
• S. antaeus (Drury), eUS, FL (CHA, HIG, IND, LAK, LEV, LIB, MARI, PUT)
  = atrolucens Casey, FL
• S. splendens (Beauvois), eUS, FL (ALA, IND)
  = cognatus Casey, FL (Kissimmee)

Tribe Dynastini

Dynastes Kirby

• D. tityus (Linnaeus), eUS, FL (ALA, LIB, VOL)

Tribe Phileurini

Phileurus Latreille

• P. truncatus (Beauvois), sUS, FL (ALA, CHA, DAD, GIL, IND, LEV, MON), Mexico, SAmer
• P. valgus (Olivier), sUS, FL (DAD, DIX, LEO, LEV, LIB, MON, PUT), Cuba, Jamaica, Mexico, CAmer
  = castaneus Haldeman
  = floridanus Casey, FL
  = sulcifer Casey, FL

Subfamily Cetoniinae

Tribe Gymnetini

Cotinis Burmeister

• C. nitida (Linnaeus), LA-IN-CT-FL (ALA, MARI)
• C. n.sp., FL (MON, Islamorada); rare; this species has been collected on only one Key; it may be a very localized endemic or it may be an immigrant.

Tribe Cetoniini

Euphoria Burmeister

• E. inda (Linnaeus), AZ-NE-CT-FL (ALA, LEV, MARI)
• E. limbalis Fall, FL endemic (ALA, DAD, LEV, MON)
• E. sepulchralis (Fabricius), TX-MO-FL (statewide); a pest of mangos in south Florida; adults feed on blossoms
• E. sp. nr. discicollis Thomson, cUS, FL (OKA); in pocket gopher mound; in R. Turnbow collection

Protaetia Burmeister

• +P. fusca (Herbst), HI, FL (BRO, DAD, PAL), immigrant from Oriental region

Stephanucha Burmeister

• S. thoracica Casey, AL, FL (ALA, GIL, LEV, MAD, OKA, WAL) (see Skelley 1991)

Tribe Cremastocheilini

Cremastocheilus Knoch

• *C. canaliculatus Kirby, IL-ON-MA, (FL?)
• C. harrisi Kirby, IL-ON-CT-FL
• C. squamulosus LeConte, NC-FL (IND); status undetermined

Tribe Trichiini

(see Howden 1968)

Trichiotinus Casey

• T. lunulatus (Fabricius), TX-NC-FL (ALA, BRO, DAD, DIX, IND, LEV, MARI)
• T. piger (Fabricius), TX-MN-VT-FL (DAD)
• T. rufobrunneus Casey, FL endemic (ALA, LEV)
• T. affinis (Gory and Percheron), eUS, FL (HAR)

Trigonopeltastes Burmeister

• T. delta (Forster), TX-MO-NJ-FL (ALA, COLU, DAD, IND, LEV, MARI, MART, MON, PUT)
• T. floridana (Casey), FL endemic (HIG, LEV, MARI, POL); at flowers of scrub palmetto; rare

Tribe Valgini

Valgus Scriba

• V. canaliculatus (Fabricius), IN-PA-NC-FL (DIX, JAC, LIB, UNI)

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